The top of molar cusps try seemingly bunoselenomorph

He’s a short and you can straight postprotocrista labially dependent. The fresh new preprotocrista, seemingly upright, suits a properly-build parastyle. The metaconule are slightly smaller which will be larger towards M3. This new premetaconulecrista therefore the fossae are simple versus neocrista. The brand new rear fossa try tilted, specifically to the M3. Out of upright to the M1, this new postmetaconulecrista will get sigmoidal on the M3, skipping brand new metacone. The newest labial cusps have been in a comparable axis having relatively straight cristae. The newest paracone labial rib is actually outstanding, but in the place of anterior groove. There isn’t any metacone labial rib. The fresh para poder-, meso-, and you can metastyles is outstanding. Brand new lingual cingulum is quite poor in order to absent including the entostyle.

The fresh filed structure of the next down premolar is common to possess derived stalk pecoran and you will top pecoran into the having a proper-arranged mesolingual conid and you will posterior cristid , . More over, the clear presence of an enamel socket on p1 excludes an attribution so you’re able to Prodremotherium or Mosaicomeryx . Even when the white teeth commonly since the bunoselenodont as with Amphitragulus elegans, the newest called specimens have very primitive have, such as for instance an extremely faster metastylid, a decreased top, a small postentocristid, and even though laterally compressed, bulged lingual cuspids toward down molars. Many of these functions exclude an association to sometimes Dremotherium or Bedenomeryx , . Furthermore, the really small size away from “Amphitragulus” quercyi is actually a determining function to your latest Oligocene ruminants out of European countries , , , .

Dremotherium cf. guthi out of Gaimersheim1: 1 remaining maxillary bone to the enamel line M1-M3 (SNSB-BSPG 1952II4855). out of Mailhat: 2 kept P4-M2 (MNHN LIM154). Babameryx engesseri from Gaimersheim1: step 3 kept fragmentary mandible that have m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and you can (c) occlusal viewpoints. The scale pub are dos cm.

Holotype called specimens information

Holotype. Jehenne (1987) enjoys appointed the new sample IPHEP LM.1969.MA.40 because holotype. That is a partial proper maxillary bones with P3-M3 (, Fig. 2), out-of Los angeles Milloque (MP29, France).

Known specimens. Mailhat MNHN LIM154 remaining P4-M2 (this really works, Fig. six.2); Gaimersheim1 SNSB-BSPG 1952II4855 leftover maxillary with M1-M3 (that it performs, Fig. six.1); SNSB-BSPG 1952II4870 correct m2.


Down permanent dentition. The reduced molar is very badly managed. not, so it large ruminant and it has a completely selenomorph trigonid. The newest cristid try elongated and cuspids is actually highest with the tooth quite pleated. The fresh new metastylid was well toned.

Higher permanent dentition. The latest P4 was defectively preserved. Every lingual part was missing. New anterolabial cone and contains a failing labial rib. The fresh anterior stylid is actually broken. The rear you’re short however, present. Zero neocrista blocks the fresh fossa. The new quadratic higher molars in occlusal take a look at sustain a proper-create metaconule quite smaller than the protocone. The fresh new cusps have become selenomorph. The latest preprotocrista plus the postmetaconulecrista are labially founded and you will some straight. The newest postprotocrista try a lot of time, very rounded, and you can labially based. It could be totally isolated (SNSB-BSPG 1952II4855) otherwise fused (MNHN LIM154) towards the premetaconulecrista. The second comes to an end at axis of your mesostyle. New posterior basin can keeps a small metaconule bend on the M2, large toward M3. This new paracone, possessing an excellent paracone rib, in addition to metacone, labially flat and concave, try aligned. The brand new mesostyle and parastyle are well set up. It mode game short articles. New mesostyle try big of M1 so you’re able to M3. The new entostyle are poor or missing. There is none a great lingual cingulum encompassing new protocone nor a metastyle developing a proper-create cuspid. However, short prior and you will rear cingula can be found.


So it ruminant is relatively state-of-the-art in becoming really-selenodont and achieving no lingual cingula during the higher molars, always contained in Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and you may Babameryx , , , [61–63], . In contrast to Bachitherium, which varieties does not have an anterior groove to your massive labial paracone rib , , . This new read fossils be more much like the state-of-the-art stem Pecora in order to the newest ancient top Pecora (such as for example Amphitragulus, Dremotherium, Bedenomeryx). The latest enamel is actually a little pleated, whereas it’s essentially easy when you look at the Prodremotherium. Moreover, how big is this type of teeth is much bigger than for the Prodremotherium on dated Quercy selections and Pech Desse . Tooth much more selenomorph than in “Amphitragulus” and Bedenomeryx , , as well as the parastyle is not that anteriorly estimated than observed towards the “Amphitragulus” feningrei , , . Studying the neodiagnosis advised by the Mennecart , these fossils become all of the exhibiting popular features of Dremotherium. The currently defined varieties inside European countries was D. feignouxi (type variety) and you may D. guthi , , . The second try a bit smaller compared to the kind kinds , much like the fossils demonstrated here. D. feignouxi merely taken place inside the Agenian angelreturn zaloguj siÄ™ (MN1-MN2; ), while D. guthi resided inside latest Oligocene (MP28-MP30; ). But not, in the lack of extra better-preserved material, a decisive assignation towards kinds is not possible.